Research in my lab deals broadly with diversity of North American fishes; taxonomic diversity, ecological diversity, and diversity of environmental adaptation. I study taxonomy, morphology, life history, and ecology of freshwater fishes.  I collect fishes in the field, I work with specimens archived in museums, and I use molecular techniques to study relationships of fishes based on DNA sequences.  Ongoing work includes studies of systematics, taxonomy, community ecology and life history of stream fishes, especially darters (Percidae: Etheostomatinae) and suckers (Catostomidae), and responses of these and other fishes to environmental degradation.

Follow links below for information on:

Training; Publications; Ongoing research; Positions; Students; Museum-related research; Human diversity efforts; Contact information; Professional fish consulting



B.S. Biology, University of New Orleans, 1979.

M.S. Biology, University of New Orleans, 1981. 

Ph.D. Zoology, University of Oklahoma, 1985.

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Recent publications

Suttkus, R.D., H.L. Bart Jr. and D. A. Etnier. 2012. A new darter of the subgenus Oligocephalus, genus Etheostoma, from southeastern Texas and southwestern Louisiana. Tulane Studies in Zoology and Botany 32(1): 6-30. [pdf]

Clements, M.D., H. L. Bart Jr. and D. L. Hurley. 2012. A different perspective on the phylogenetic relationships of the Moxostomatini (Cypriniformes: Catostomidae) based on cytochrome-b and Growth Hormone intron sequences. Molecular Phylogenetics and Evolution 63:159–167. [pdf]

Doosey, M.H. and H. L. Bart Jr. 2011. Morphological variation of the palatal organ and chewing pad of catostomidae (Teleostei: Cypriniformes). J. Morphology 272: 1092-1108. [pdf]

Tang, K. L. M.K. Agnew, W-J. Chen, M.V. Hirt, M.E. Raley, T. Sado, L.M. Schneider, L. Yang, H. L. Bart Jr., S. He, H. Liu, M, Miya, K, Saitoh, A.M. Simons, R.M. Wood, R.L. Mayden. 2011. Phylogeny of the gudgeons (Teleostei: Cyprinidae: Gobioninae). Molecular Phylogenetics and Evolution 61: 103–124. [pdf]

Cashner, M.F., K.R. Piller, H.L. Bart Jr. 2011. Phylogenetic relationships of the North American cyprinid subgenus Hydrophlox. Molecular Phylogenetics and Evolution 59: 725–735. [pdf]

Albert, J.S., H.L. Bart Jr. and Roberto E. Reis. 2011. Species richness and cladal diversity. pp 89-104 In: J.S. Albert and R.E. Reis (Eds.) Historical Biogeography of Neotropical Freshwaters. University of California Press. [pdf]

Saitoh, K., T. Sado, M.H. Doosey, H.L. Bart Jr., J.G. Inoue, M. Nishida, R.L. Mayden, and M. Miya. 2011. Evidence from mitochondrial genomics supports the lower Mesozoic of South Asia as the time and place of basal divergence of cypriniform fishes (Actinopterygii: Ostariophysi). Zoological Journal of the Linnean Society 161:633–662. [pdf]

Cashner, M.F. and H. L. Bart Jr. 2010. Reproductive Ecology of Nest Associates: Use of RFLPs to Identify Cyprinid Eggs. Copeia 2010(4): 554-557.[pdf]

Bart, H.L. Jr., M.D. Clements, R. E. Blanton, K. R. Piller, D. L. Hurley. 2010. Discordant molecular and morphological evolution in buffalofishes (Actinopterygii: Catostomidae). Molecular Phylogenetics and Evolution 56 (2010) 808–820. [pdf]

Bart, H.L. Jr., P.C. Reneau, M.H. Doosey, C.D. Bell. 2010. Evolutionary divergence of duplicate copies of the growth hormone gene in suckers (Actinopterygii: Catostomidae). International Journal of Molecular Sciences 11: 1090-1102. [pdf]

Doosey, M. H., H. L. Bart, Jr., K. Saitoh, and M. Miya. 2010. Phylogenetic relationships of Catostomid Fishes (Actinopterygii: Cypriniformes) based on Mitochondrial ND4/ND5 Gene Sequences. Molecular Phylogenetics and Evolution 54 (2010) 1028–1034. [pdf]

Chen, Y., S. Huang, H. Chen, and H. L. Bart, Jr. 2010. Joint feature selection and classification for taxonomic problems within fish species complexes. Pattern Analysis and Applications 13 (2): 23–34. [pdf]

Piller, K. R. and H. L. Bart, Jr. 2009. Incomplete sampling, outgroups, and phylogenetic inaccuracy: A case study of the Greenside Darter complex (Percidae: Etheostoma blennioides). Molecular Phylogenetics and Evolution 53 (2009) 340–344. [pdf]

Mayden, R.L. W-J. Chen, H.L. Bart, Jr., M.H. Doosey, A.M. Simons, K.L. Tang, R.M. Wood, M.K. Agnew, L. Yang, M.V. Hirt, M.D. Clements, K. Saitoh, T. Sado, M. Miya, M. Nishida. 2009. Reconstructing the Phylogenetic Relationships of the Earth’s 1 Most Diverse Clade of Freshwater Fishes - Order Cypriniformes (Actinopterygii: Ostariophysi): A Case Study Using Multiple Nuclear Loci and the Mitochondrial Genome. Molecular Phylogenetics and Evolution 51:500-514. [pdf]

Chen, Y, X. Dang, H. Peng, and H. L. Bart, Jr. 2009. Outlier Detection with the Kernelized Spatial Depth Function.  IEEE Transactions on Pattern Analysis and Machine Intelligence, vol. 31, no. 2, pp:288-305. [pdf]

Mayden, R.L., K.L. Tang, R. M. Wood, W-J, Chen, M. K. Agnew, K.W. Conway, L. Yang, A.M. Simons, H.L. Bart, Jr., P.M. Harris, J. LI, X. Wang, K. Saitoh, S. He, H. Liu, Y. Chen, M. Nishida, M. MIYA. 2008. Inferring the Tree of Life of the order Cypriniformes, the earth’s most diverse clade of freshwater fishes: Implications of varied taxon and character sampling. Journal of Systematics and Evolution 46 (3): 424–438. [pdf]

Chen, H., H.L. Bart, Jr. and S. Huang. 2008. Integrated Feature Selection and Clustering for Taxonomic Problems within Fish Species Complexes. Journal of Multimedia 3:10-17. [pdf]

Piller, K.R., H. L. Bart, Jr. and D. L. Hurley. 2008. Phylogeography of the greenside darter complex, Etheostoma blennioides (Teleostomi: Percidae): A wide-ranging polytypic taxon. Molecular Phylogenetics and Evolution 46: 974–985. [pdf]

Chen, H., S. Huang and H. L. Bart, Jr.  2006. Taxonomy of fish species complexes: a role for multimedia information. Pp 475-479 In: Proceedings of the 8th IEEE, International Workshop on Multi Media Signal Processing, Victoria, British Columbia. [pdf]

Fentress, J. A., S. L. Steele, H. L. Bart, Jr. and Ann Oliver Cheek. 2006. Reproductive Disruption in Wild Longear Sunfish (Lepomis megalotis) Exposed to Kraft Mill Effluent. Environmental Health Perspectives 114: 40-45. [pdf]

Chen, Y., H.L. Bart, Jr., and F. Teng. 2005. A Content-Based Image Retrieval System for Fish Taxonomy. 7th ACM SIGMM International Workshop on Multimedia Information Retrieval, pp. 237-244, Singapore, 2005. [pdf]

Chen, Y., H. L. Bart, Jr., S. Huang, and H. Chen. 2005. A computational framework for taxonomic research: diagnosing body shape within fish species complexes. Proc. IEEE Int’l Conf. on Data Mining, pp. 593-596. [pdf]

Chen, Y., H. L. Bart, Jr., X. Dang, and H. Peng.  2005. Depth-Based Novelty Detection and its Application to Taxonomic Research, pp: 113-122 In: Proceedings 7th IEEE International Conference on Data Mining (ICDM). [pdf]

Freeman, M.J., E.R. Irwin, N.M. Burkhead, B.J. Freeman, H.L. Bart, Jr. Status and conservation of the Fish Fauna of the Alabama River System. American Fisheries Society Symposium 45:557–585. [pdf]

Watanabe, K. H. H, Lin, H.L. Bart, Jr., P.J. Martinat, J. C. Means, M.L. Kunas, D.A. Grimm. 2005. Bayesian estimation of kinetic rate constants in a food-web model of polycyclic aromatic hybrocarbon accumulation. Ecological Modelling 181: 229-246. [pdf]

Greenfield, D. I. and H.L. Bart, Jr.  2005. Fish community dynamics over a sixteen-year collection period from a blackwater stream receiving paper mill effluents. Hydrobiologia 534: 81-90. [pdf]

Page, L. M., H.L. Bart, Jr., R. Beaman, L. Bohs, L. T. Deck, V. A. Funk, D. Lipscomb, M. A. Mares, L. A. Prather, J. Stevenson, Q. D. Wheeler, J. B. Woolley, D. W. Stevenson. 2005. LINNE: Legacy Infrastructure Network for Natural Environments. Illinois Natural History Survey, Champaign, IL []

Bart, H. L., Jr., R. D. Suttkus, J. Lyons and N. Mercado-Silva. 2004. A preliminary analysis of the taxonomic status of Ictiobus meridionalis (Günther). pp 47-61 In: M.L. Lozano-Vilano and A.J. Contreras-Balderas (eds), Homenaje al Doctor Andres Resendez Medina, Universidad Autonoma de Nuevo Leon, Monterrey, Mexico. [pdf

Tipton, J. A., H. L. Bart, Jr. and K. R. Piller. 2004. Geomorphic disturbance and its impact on darter (Teleostomi: Percidae) distribution and abundance in the Pearl River drainage, Mississippi. Hydrobiologia 527: 49–61. [pdf]

Piller, K.R., H.L. Bart, Jr. and J. A. Tipton. 2004. Decline of the frecklebelly madtom in the Pearl River Basin based on Historical and Contemporary Surveys. Transactions of the American Fisheries Society 133: 1004-1013. [pdf]

Clements, M.D., H.L. Bart, Jr. and D.L. Hurley. 2004 .Isolation and characterization of two distinct Growth Hormone cDNAs from the tetraploid smallmouth buffalofish (Ictiobus bubalus)", General and Comparative Endocrinology 136: 411-418. [pdf]

Nickum, J.G., H. L. Bart, Jr., P.R. Bowser, I. E. Greer, C. Hubbs, J. A. Jenkins, J. R. MacMillan, J. W. Rachlin, J. D. Rose, P. W. Sorensen, J. R. Tomasso. 2004. Guidelines for the use of fishes in research. American Fisheries Society, Bethesda MD ( 

Piller, K.R. H.L. Bart, Jr. and J.A. Tipton. 2003. Spawning of the black buffalo, Ictiobus niger (Cypriniformes: Catostomidae). Ichthyological Explorations of Freshwaters 14:145-150. [pdf]

Machado, M.D., D.C. Heins, and H.L. Bart, Jr. 2002 Microgeographic variaion in ovum size of the blacktail shiner, Cyprinella venusta Girard, in relation to streamflow. Ecology of Freshwater Fish. 11:11-19. [pdf]

Suttkus, R.D. and H.L. Bart, Jr. 2002. A preliminary analysis of the river carpsucker, Carpiodes carpio, in the southern portion of its range. In: L. Lozano (ed.) The professional career of Salvador Contreras-Balderas, Universidad Autonoma de Nuevo Leon. [pdf]

Watanabe, K and H. L. Bart, Jr. 2001 Comments on "Model of biota-sediment accumulation factor for polycyclic aromatic hydrocarbons". Envir. Tox. And Chem.  20(9): 1867-1869.

Piller, K.R., H.L. Bart, Jr. and C.A. Walser. 2001. Morphological variation of the redfin darter, Etheostoma whipplei, with comments on the status of the subspecific populations. Copeia 2001 (3):802-807. [pdf]

Bart, H.L., Jr.  2000. Fish diversity in a heavily industrialized reach of the lower Mississippi River, pp 203-218 In: C. Colten (ed.) Centuries of Change: human transformation of the lower Mississippi River. University of Pittsburgh Press. [pdf]

Walser, C. A., B. Falterman, and H.L. Bart, Jr.  2000. Impact of introduction of the rough shiner (Notropis baileyi) on the native fish community of the Chattachoochee River system. American Midland Naturalist 144:393-405. [pdf]

Warren, M. L. Jr., B. M. Burr, S. J. Walsh, H. L. Bart, Jr., R. C. Cashner, D. A. Etnier, B.J. Freeman, B. R. Kuhajda, R. L. Mayden, H. W. Robison, S. T. Ross, and W. C. Starnes. 2000. Diversity, distribution, and conservation status of the native freshwater fishes of the southern United States. Fisheries 25(10): 7-31.

Complete publication list

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Ongoing research projects 

Systematics of Fishes of Subfamily Ictiobinae

Systematics and Biodiversity of Cypriniformes

Spatial network of bioaccumulation in food webs

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Current Positions

Director, Tulane University Biodiversity Research Institute (2011-present)

Director, Tulane University Museum of Natural History (1993-2011)

Curator of Fishes Royal D. Suttkus Fish Collection (1992-present)

Professor of Ecology and Evolutionary Biology, Tulane University (2007-present)      

Associate Professor of Ecology and Evolutionary Biology, Tulane University (1996-2007)                      

Editor of Tulane Studies in Zoology and Botany and Occasional Papers Tulane University Museum of Natural History (1994-present). 


Previous positions   

Assistant Professor, Ecology and Evolutionary Biology, Tulane University (1992-95)

Curator of Fishes, Auburn University Museum Fish Collection (1988-91)

Assistant Professor, Zoology and Wildlife Science, Auburn University (1988-91)

Postdoctoral Research Associate, Ecology, Ethology and Evolution, University of Illinois (1986-88), Lawrence M. Page (Mentor)

Visiting Assistant Professor, Ecology, Ethology and Evolution, University of Illinois (1995-88)

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Human diversity efforts

I coordinated Science Technology Engineering and Mathematics (STEM) Minority training programs at the undergraduate and graduate levels at Tulane from 1996-2012.  I served as Campus Coordinator for the Tulane Campus Program in the Louis Stokes Louisiana Alliance for Minority Participation (1996-2010) and Co-PI and Project Director for the Graduate Alliance for Education in Louisiana (STEM minority doctoral training program) from 2002-2012.  I actively involve minorities in my research in ecology and systematics and work to recruit minorities to careers in these areas.

Tulane LS-LAMP Program

Graduate Alliance for Education in Louisiana

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Collection-related research (follow links below to more detailed descriptions of projects):

I direct the efforts of curatorial staff at TUBRI to care for and facilitate use of the Royal D. Suttkus Fish Collection.  Together with Nelson Rios and other TUBRI staff, we design software tool for automated georeferencing of electronic collection location records.  We also manage the Fishnet2 network of fish collection databases. I also participate in or direct externally-funded research on fish systematics, taxonomy, specimen imaging and analysis.


Fishnet II (Collaborative)

Curatorial improvements to the Tulane Fish Collection

Tulane University Biodiversity Research Institute

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Current students (and their research)

Ray Schmidt

Justin Mann


Former students

Mollie Cashner, Ph.D. Biology, Tulane University, 2010

Michael Doosey, Ph.D. Biology, Tulane University, 2009

Mark Clements, Ph.D. Biology, Tulane University, 2008 

Rebecca Johansen, Ph.D. Biology, Tulane University, 2007

Jeanine Burse, Ph.D., Biology, Tulane University, 2004

Kyle Piller, Ph.D., Biology, Tulane University, 2001

Christoph Walser, Ph.D., Biology, Tulane University, 1996

Myra Kunas, M.S., Biology, Tulane University, 2003

Jason Tipton, M.S., Biology, Tulane University, 2002

Joseph Buckley, M.S., Biology, Tulane University, 1995

Dianne Greenfield, M.S., Biology, Tulane University, 1995 

Andrea Brooks, M.S. Zoology and Wildlife Science, Auburn University, 1993

Jeffrey Harbaugh, M.S., Zoology and Wildlife Science, Auburn University, 1994

Postdoctoral Research Associates

Chuck Bell, 2005-2006

Paulette Reneau, 2008-2011

Michael Doosey, 2012-present


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Contact Information

Tulane University Biodiversity Research Institute
Building A-3, Wild Boar Rd.
Belle Chasse, LA 70037
E-mail address:
Voice: 504-394-1711
Fax: 504-394-5045

Department of Ecology and Evolutionary Biology
Tulane University
Room 310, Dinwiddie Hall,
6823 St. Charles Ave.
New Orleans LA 70118
504-862-8706 (fax)

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